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J Biol Chem 254:51845190 Osawa HG, Stacey W, Gassmann S (2006) OPT1 and AtOPT4 function as proton-coupled oligopeptide transporters with broad but distinct substrate specificities

Not only surface markers but also certain histologically stained intracellular components can be regarded as activation markers, since activated eosinophils undergo degranulation and release granule proteins detectable by specific immunostaining