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SS-31 Research Peptide Discovery Context 2013 Weill Cornell Medical College: SS-31 research peptide (Elamipretide) was characterized during investigations into mitochondria-targeted tetrapeptides designed to study: cardiolipin-binding mechanisms mitochondrial-protein interaction networks oxidative-stress modulation electron-transport efficiency bioenergetic resilience under metabolic strain Subsequent research established SS-31 research peptide as a widely studied substrate for exploring mitochondrial membrane stabilization , protein-interaction remodeling , and bioenergetic regulation in controlled laboratory systems.